Reproduced locally.
./out/Debug/browser_tests \
--gtest_filter="OmniboxPopupContentsViewTest.ThemeIntegration*" \
--enable-features=Mash
> Cr-Commit-Position: refs/heads/master@{#556454} good
> Cr-Commit-Position: refs/heads/master@{#556646} bad
So yeap, crrev.com/556646 was the cause.
Comment 1 by ksakamoto@chromium.org
, May 8 2018